The wide deployment of vehicle automation and communication systems (VACS) in the next decade is expected to influence traffic performance on freeways. Apart from safety and comfort, one of the goals is the alleviation of traffic congestion which is a major and challenging problem for modern societies. The paper investigates the combined use of two feedback control strategies utilizing VACS at different penetration rates, aiming to maximize throughput at bottleneck locations. The first control strategy employs mainstream traffic flow control using appropriate variable speed limits as an actuator. The second control strategy delivers appropriate lane-changing actions to selected connected vehicles using a feedback-feedforward control law. Investigations of the proposed integrated scheme have been conducted using a microscopic simulation model for a hypothetical freeway featuring a lane-drop bottleneck. The results demonstrate significant improvements even for low penetration rates of connected vehicles.